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["Fok Evolutie"]
["\"Niets in de biologie heeft betekenis behalve in het licht van de evolutie\" (Theodosius Dobzhansky, 1973)"]
[]
["Kippen zijn stiekem erg interessante dieren vanuit een evolutionair perspectief. Wist je dat kippen relatief weinig genetische veranderingen ondergaan hebben sinds ze zich afsplitsten van gevederde dinosaurussen?\nUit een studie bleek dat van de 21 onderzochte vogelsoorten, het DNA van de kip het meest op het (gereconstueerde) DNA van een dinosaurier lijkt (Romanov et al, 2014).\nDe komende lessen ga je onderzoeken hoe (een gedeelte van) de evolutie van een kip had kunnen plaatsvinden! We starten met een klein stukje herhaling van het thema \"erfelijkheid\" door eerst een kip te fokken."]
[]
[null,"Dit is wat enkele leerlingen antwoordden op de vraag \"wat vond je van deze opdracht?\""]
[null,"\"Goede inzichtopdrachten die bij het boek passen\""]
[null,"\"Ik vond het grappig / leuk om te doen, het voelde niet echt als huiswerk.\""]
[null,"\"Ja best leuk eigenlijk, omdat het heel anders was dan andere projecten en ik vond het wel fijn werken in excel.\""]
[null,"\"Heel leuk. Het was behoorlijk wat moeite moet ik bekennen. maar ik vond het erg interessant en leuk om te doen.\""]
[null,"\"heel erg leuk en origineel\""]
[null,"\"Ik vond het een leuke opdracht. Je moest echt goed nadenken, want het was soms best lastig.\""]
[null,"\"Erg leuk, soms wel met wat pittige vragen, maar wel leuk om er over na te denken. Ook vond ik het goed dat u de opdrachten besprak in de les, zodat je je antwoorden kon vergelijken.\""]

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["Fok je ideale kip"]
["\"Niets in de biologie heeft betekenis behalve in het licht van de evolutie\" (Theodosius Dobzhansky, 1973)"]
["Vul alleen gegevens in, in de groene velden. Wanneer je gegevens verandert in de andere cellen loop je kans dat de formules niet meer correct functioneren en daarmee de opdracht niet meer goed werkt."]
["1: vul onderstaande gegevens in"]
["Naam:"]
["klas:"]
["datum:"]
[null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,0.12280271561301637]
["2: Je gaat  je ‘ideale’ kip proberen te fokken. Om kippen te kunnen fokken, moet je het een en ander weten over genen en kruisingen. Zorg ervoor dat je de volgende begrippen kent: gen, allel, dominant, recessief, homozygoot en heterozygoot. Deze opdracht heb je al eerder gedaan in de klas met kaartjes in een groepje. Omdat het thema \"Evolutie\"  voortborduurt op de thema's \"Erfelijkheid\" en \"Soorten en Populaties\" is het belangrijk om met deze opdracht het geheugen nog eens op te frissen. Geen paniek; doordat de opdracht nu digitaal is gemaakt, kun je erg snel je resultaten kopiëren en hergebruiken.  De resultaten die je in deze opdracht krijgt, heb je nodig om de vervolg opdrachten de komende weken te kunnen maken. Start met het omschrijven van onderstaande begrippen."]
["gen:"]
["allel:"]
["dominant:"]
["recessief:"]
["homozygoot:"]
["heterozygoot:"]
[]
["3: Klik op de knop \"bekijk opdracht \"om te zien of jij een vleeskip, legkip of sierkip gaat fokken!"]
["opdracht:"]
[]
["4: Je kunt bij het fokken letten op vijf verschillende genen (A t/m E). Van ieder gen komen twee varianten voor: een dominant allel (aangegeven met een hoofdletter) en een recessief allel (aangegeven met een kleine letter). In het schema hieronder kun je aflezen voor welke eigenschappen deze allelen coderen."]
[0.68432866279856719]
["Allel A/a ","Dit gen codeert voor een groeihormoon, een eiwit dat onder andere zorgt voor de aanmaak van spierweefsel. De volgende allelen en fenotypen komen voor: "]
[null,"A: De kip heeft na 6 weken een lichaamsgewicht van 2,5 kg. Door de snelle groei heeft de kip gezondheidsproblemen en leeft niet langer dan een half jaar. "]
[null,"a: De kip bereikt na 14 weken een lichaamsgewicht van 2,5 kg. De kip kan onder normale omstandigheden vijf jaar of langer in leven blijven."]
[]
["Allel B/b","Dit gen codeert voor een hormoon dat invloed heeft op het sociaal gedrag van de kip. De volgende allelen en fenotypen komen voor: "]
[null,"B: De kip vertoont agressief gedrag (pikken) richting soortgenoten, met name wanneer de kippen dicht op elkaar zitten of wanneer de eigen kuikens worden aangevallen."]
[null,"b: De kip is relatief passief richting soortgenoten; de eigen kuikens worden nauwelijks beschermd tegen aanvallen van andere kippen."]
[]
["Allel C/c ","Dit gen codeert voor een enzym dat een rol speelt bij de pigmentatie (kleuring) van de veren. De volgende allelen en fenotypen komen voor:"]
[null,"C: De kip heeft egaal witte veren. "]
[null,"c: De kip heeft zwart-wit gespikkelde veren."]
[]
["Allel D/d ","Dit gen codeert voor een eiwit dat invloed heeft op de hoeveelheid energie die een kip investeert in voortplanting. De volgende allelen en fenotypen komen voor: "]
[null,"D: De kip investeert veel energie in voortplanting, en produceert gemiddeld 200 eieren per jaar. Hierdoor is de lichaamsgrootte van het dier beperkt."]
[null,"d: De kip investeert minder energie in voorplanting, en kan per jaar hooguit 50 eieren produceren. De eierproductie van het dier gaat niet ten koste van de lichaamsgroei."]
[]
["Allel E/e ","Dit gen codeert voor een enzym dat invloed heeft op de veergroei op de kop van de kip. De volgende allelen en fenotypen komen voor:"]
[null,"E: De kip heeft een groot aantal kuifveren op zijn kop. "]
[null,"e: De kip heeft geen kuifveren op zijn kop."]
[]
["5: Er zijn vijf genen waar je op kan letten (A t/m E). Lees hierboven de informatie over deze vijf kippengenen goed door. Geef aan op welke drie genen jullie het meest gaan letten en aan welk genotype je ideale kip voor deze drie genen heeft. Zet je antwoorden in de tabel hieronder."]
["Allel A/a ","mijn ideale genotype voor gen A/a (groeifactor) is:"]
["Allel B/b","mijn ideale genotype voor gen B/b (sociaal gedrag) is:"]
["Allel C/c ","mijn ideale genotype voor gen C/c (voortplanting) is:"]
["Allel D/d ","mijn ideale genotype voor gen D/d (kuif veren) is:"]
["Allel E/e ","mijn ideale genotype voor gen E/e (kuif) is:"]
[]
[null,"Hanen",null,null,null,null,null,null,null,"Hennen"]
[null,"Bob",null,"Dirk",null,"Otto",null,null,null,"Anne",null,"Christina",null,"Puk"]
["gen1","a","a","A","a","a","a",null,null,"a","a","a","a","a","a"]
["gen2","B","B","B","B","B","b",null,null,"B","b","B","B","B","B"]
["gen3","C","c","C","C","C","c",null,null,"C","C","C","C","C","C"]
["gen4","d","d","d","d","d","d",null,null,"d","d","D","d","d","d"]
["gen5","e","e","E","E","e","e",null,null,"e","e","e","e","E","e"]
[]
[]
["6: Deze kippen vormen de P generatie. Bekijk de genotypen van deze kippen. Bepaal welke twee kippen (één haan en één hen) je met elkaar wilt laten voortplanten. Kopieer de naam en het genotype van de twee kippen waarmee je gaat starten in de groene velden hieronder "]
["P) We kruisen door met deze twee kippen:"]
[]
[null,null,null,null,"haan:","Naam",null,null,"hen:","Naam"]
[null,null,null,null,null,"A/a","A/a",null,null,"A/a","A/a"]
[null,null,null,null,null,"B/b","B/b",null,null,"B/b","B/b"]
[null,null,null,null,null,"C/c","C/c",null,null,"C/c","C/c"]
[null,null,null,null,null,"D/d","D/d",null,null,"D/d","D/d"]
[null,null,null,null,null,"E/e","E/e",null,null,"E/e","E/e"]
[]
[null,null,null,null,null,{"$f":"F61","v":"Naam"},null,0,null,{"$f":"J61","v":"Naam"},null,0.42456969103358311]
[null,null,null,null,"gen1",{"$f":"F62","v":"A/a"},{"$f":"G62","v":"A/a"},0,null,{"$f":"J62","v":"A/a"},{"$f":"K62","v":"A/a"},0]
[null,null,null,null,"gen2",{"$f":"F63","v":"B/b"},"B/b",0,null,{"$f":"J63","v":"B/b"},"B/b",0]
[null,null,null,null,"gen3","C/c","C/c",0,null,"C/c","C/c",0]
[null,null,null,null,"gen4","D/d","D/d",0,null,{"$f":"J65","v":"D/d"},{"$f":"K65","v":"D/d"},0]
[null,null,null,null,"gen5","E/e","E/e",0,null,{"$f":"J66","v":"E/e"},"E/e",0]
[]
[null,null,null,null,null,"zaadcel:",null,"Embryo",null,null,"eicel:"]
[null,null,null,null,"gen1",{"$f":"IF(H69<0.5,F69,G69)","v":"A/a"},null,"A","a",null,{"$f":"IF(L69<0.5,J69,K69)","v":"A/a"}]
[null,null,null,null,"gen2",{"$f":"IF(H70<0.5,F70,G70)","v":"B/b"},null,"B","B",null,{"$f":"IF(L70<0.5,J70,K70)","v":"B/b"}]
[null,null,null,null,"gen3",{"$f":"IF(H71<0.5,F71,G71)","v":"C/c"},null,"C","C",null,{"$f":"IF(L71<0.5,J71,K71)","v":"C/c"}]
[null,null,null,null,"gen4","D/d",null,"d","d",null,{"$f":"IF(L72<0.5,J72,K72)","v":"D/d"}]
[null,null,null,null,"gen5","E/e",null,"E","E",null,{"$f":"IF(L73<0.5,J73,K73)","v":"E/e"}]
[]
[]
["7: Genereer de F1 door op de knop \"kruis dieren te klikken\"."]
["Noteer de zes genotypen van de F1 die is ontstaan door de dieren te kruisen en het genotype van het embryo te kopieren naar onderstaande tabel. Geef ieder nieuw dier een naam"]
[]
[]
[]
[null,"Hanen",null,null,null,null,null,null,null,"Hennen"]
["F1","naam 1",null,"naam 2",null,"naam3",null,null,null,"naam 1",null,"naam 2",null,"naam 3"]
["gen1"]
["gen2"]
["gen3"]
["gen4"]
["gen5"]
[]
["8: Kopieer de naam en het genotype van de 2 kippen uit de F1 waarmee je doorkruist in de groene velden hieronder. We kruisen door met deze twee kippen:"]
[]
[null,null,null,null,"haan:","Naam",null,null,"hen:","Naam"]
[null,null,null,null,null,"A/a","A/a",null,null,"A/a","A/a"]
[null,null,null,null,null,"B/b","B/b",null,null,"B/b","B/b"]
[null,null,null,null,null,"C/c","C/c",null,null,"C/c","C/c"]
[null,null,null,null,null,"D/d","D/d",null,null,"D/d","D/d"]
[null,null,null,null,null,"E/e","E/e",null,null,"E/e","E/e"]
[]
[null,null,null,null,null,{"$f":"F98","v":"Naam"},null,8.0637546884134492E-2,null,{"$f":"J98","v":"Naam"},null,0.45743929779580361]
[null,null,null,null,"gen1",{"$f":"F99","v":"A/a"},{"$f":"G99","v":"A/a"},0,null,{"$f":"J99","v":"A/a"},{"$f":"K99","v":"A/a"},0]
[null,null,null,null,"gen2",{"$f":"F100","v":"B/b"},"B/b",0,null,{"$f":"J100","v":"B/b"},"B/b",0]
[null,null,null,null,"gen3",{"$f":"F101","v":"C/c"},"C/c",0,null,{"$f":"J101","v":"C/c"},"C/c",0]
[null,null,null,null,"gen4",{"$f":"F102","v":"D/d"},"D/d",0,null,{"$f":"J102","v":"D/d"},{"$f":"K102","v":"D/d"},0]
[null,null,null,null,"gen5",{"$f":"F103","v":"E/e"},"E/e",0,null,{"$f":"J103","v":"E/e"},"E/e",0]
[]
[null,null,null,null,null,"zaadcel:",null,"Embryo",null,null,"eicel:"]
[null,null,null,null,"gen1",{"$f":"IF(H106<0.5,F106,G106)","v":"A/a"},null,"A/a","A/a",null,{"$f":"IF(L106<0.5,J106,K106)","v":"A/a"}]
[null,null,null,null,"gen2",{"$f":"IF(H107<0.5,F107,G107)","v":"B/b"},null,"B/b","B/b",null,{"$f":"IF(L107<0.5,J107,K107)","v":"B/b"}]
[null,null,null,null,"gen3",{"$f":"IF(H108<0.5,F108,G108)","v":"C/c"},null,"C/c","C/c",null,{"$f":"IF(L108<0.5,J108,K108)","v":"C/c"}]
[null,null,null,null,"gen4","D/d",null,"D/d","D/d",null,{"$f":"IF(L109<0.5,J109,K109)","v":"D/d"}]
[null,null,null,null,"gen5","E/e",null,"E/e","E/e",null,{"$f":"IF(L110<0.5,J110,K110)","v":"E/e"}]
[]
[]
["9: Genereer de F2 door op de knop \"kruis dieren te klikken F1\"."]
["Noteer de zes genotypen van de F2 die is ontstaan door de dieren te kruisen en het genotype van het embryo te kopieren naar onderstaande tabel. Geef ieder nieuw dier een naam"]
[]
[]
[]
[null,"Hanen",null,null,null,null,null,null,null,"Hennen"]
["F2","naam 1",null,"naam 2",null,"naam3",null,null,null,"naam 1",null,"naam 2",null,"naam 3"]
["gen1"]
["gen2"]
["gen3"]
["gen4"]
["gen5"]
[]
["10: Kopieer de naam en het genotype van de 2 kippen uit de F2 waarmee je doorkruist in de groene velden hieronder "]
["We kruisen door met deze twee kippen:"]
[]
[null,null,null,null,"haan:","Naam",null,null,"hen:","Naam"]
[null,null,null,null,null,"A/a","A/a",null,null,"A/a","A/a"]
[null,null,null,null,null,"B/b","B/b",null,null,"B/b","B/b"]
[null,null,null,null,null,"C/c","C/c",null,null,"C/c","C/c"]
[null,null,null,null,null,"D/d","D/d",null,null,"D/d","D/d"]
[null,null,null,null,null,"E/e","E/e",null,null,"E/e","E/e"]
[]
[null,null,null,null,null,{"$f":"F136","v":"Naam"},null,null,null,{"$f":"J136","v":"Naam"}]
[null,null,null,null,"gen1",{"$f":"F137","v":"A/a"},{"$f":"G137","v":"A/a"},0,null,{"$f":"J137","v":"A/a"},{"$f":"K137","v":"A/a"},0]
[null,null,null,null,"gen2",{"$f":"F138","v":"B/b"},"B/b",0,null,{"$f":"J138","v":"B/b"},"B/b",0]
[null,null,null,null,"gen3",{"$f":"F139","v":"C/c"},"C/c",0,null,{"$f":"J139","v":"C/c"},"C/c",0]
[null,null,null,null,"gen4",{"$f":"F140","v":"D/d"},"D/d",0,null,{"$f":"J140","v":"D/d"},{"$f":"K140","v":"D/d"},0]
[null,null,null,null,"gen5",{"$f":"F141","v":"E/e"},"E/e",0,null,{"$f":"J141","v":"E/e"},"E/e",0]
[]
[null,null,null,null,null,"zaadcel:",null,"Embryo",null,null,"eicel:"]
[null,null,null,null,"gen1",{"$f":"IF(H144<0.5,F144,G144)","v":"A/a"},null,"A/a","A/a",null,{"$f":"IF(L144<0.5,J144,K144)","v":"A/a"}]
[null,null,null,null,"gen2",{"$f":"IF(H145<0.5,F145,G145)","v":"B/b"},null,"B/b","B/b",null,{"$f":"IF(L145<0.5,J145,K145)","v":"B/b"}]
[null,null,null,null,"gen3",{"$f":"IF(H146<0.5,F146,G146)","v":"C/c"},null,"C/c","C/c",null,{"$f":"IF(L146<0.5,J146,K146)","v":"C/c"}]
[null,null,null,null,"gen4","D/d",null,"D/d","D/d",null,{"$f":"IF(L147<0.5,J147,K147)","v":"D/d"}]
[null,null,null,null,"gen5","E/e",null,"E/e","E/e",null,{"$f":"IF(L148<0.5,J148,K148)","v":"E/e"}]
[]
[]
["11: Genereer de F3 door op de knop \"kruis dieren te klikken F2\"."]
["Noteer de zes genotypen van de F3 die is ontstaan door de dieren te kruisen en het genotype van het embryo te kopieren naar onderstaande tabel. Geef ieder nieuw dier een naam"]
[]
[]
[]
[null,"Hanen",null,null,null,null,null,null,null,"Hennen"]
["F3","naam 1",null,"naam 2",null,"naam3",null,null,null,"naam 4",null,"naam 5",null,"naam 6"]
["gen1"]
["gen2"]
["gen3"]
["gen4"]
["gen5"]
[]
["12: Kopieer de naam en het genotype van de 2 kippen uit de F3 waarmee je doorkruist in de groene velden hieronder "]
["We kruisen door met deze twee kippen:"]
[]
[null,null,null,null,"haan:","Naam",null,null,"hen:","Naam"]
[null,null,null,null,null,"A/a","A/a",null,null,"A/a","A/a"]
[null,null,null,null,null,"B/b","B/b",null,null,"B/b","B/b"]
[null,null,null,null,null,"C/c","C/c",null,null,"C/c","C/c"]
[null,null,null,null,null,"D/d","D/d",null,null,"D/d","D/d"]
[null,null,null,null,null,"E/e","E/e",null,null,"E/e","E/e"]
[]
[null,null,null,null,null,{"$f":"F174","v":"Naam"},null,null,null,{"$f":"J174","v":"Naam"}]
[null,null,null,null,"gen1",{"$f":"F175","v":"A/a"},{"$f":"G175","v":"A/a"},0,null,{"$f":"J175","v":"A/a"},{"$f":"K175","v":"A/a"},0]
[null,null,null,null,"gen2",{"$f":"F176","v":"B/b"},"B/b",0,null,{"$f":"J176","v":"B/b"},"B/b",0]
[null,null,null,null,"gen3",{"$f":"F177","v":"C/c"},"C/c",0,null,{"$f":"J177","v":"C/c"},"C/c",0]
[null,null,null,null,"gen4",{"$f":"F178","v":"D/d"},"D/d",0,null,{"$f":"J178","v":"D/d"},{"$f":"K178","v":"D/d"},0]
[null,null,null,null,"gen5",{"$f":"F179","v":"E/e"},"E/e",0,null,{"$f":"J179","v":"E/e"},"E/e",0]
[]
[null,null,null,null,null,"zaadcel:",null,"Embryo",null,null,"eicel:"]
[null,null,null,null,"gen1",{"$f":"IF(H182<0.5,F182,G182)","v":"A/a"},null,"A","a",null,{"$f":"IF(L182<0.5,J182,K182)","v":"A/a"}]
[null,null,null,null,"gen2",{"$f":"IF(H183<0.5,F183,G183)","v":"B/b"},null,"B","B",null,{"$f":"IF(L183<0.5,J183,K183)","v":"B/b"}]
[null,null,null,null,"gen3",{"$f":"IF(H184<0.5,F184,G184)","v":"C/c"},null,"C","C",null,{"$f":"IF(L184<0.5,J184,K184)","v":"C/c"}]
[null,null,null,null,"gen4","D/d",null,"d","d",null,{"$f":"IF(L185<0.5,J185,K185)","v":"D/d"}]
[null,null,null,null,"gen5","E/e",null,"E","e",null,{"$f":"IF(L186<0.5,J186,K186)","v":"E/e"}]
[]
[]
["13: Genereer de F4 door op de knop \"kruis dieren te klikken F3\"."]
["Noteer de zes genotypen van de F4 die is ontstaan door de dieren te kruisen en het genotype van het embryo te kopieren naar onderstaande tabel. Geef ieder nieuw dier een naam"]
[]
[]
[]
[null,"Hanen",null,null,null,null,null,null,null,"Hennen"]
["F4","naam 1",null,"naam 2",null,"naam3",null,null,null,"naam 4",null,"naam 5",null,"naam 6"]
["gen1"]
["gen2"]
["gen3"]
["gen4"]
["gen5"]
[]
["14: Bekijk de kippen in je F4. Hebben je kippen het genotype dat je wilde? Met andere woorden, is het gelukt om je ideale kip te fokken? "]
[]
[]
["Zo nee, wat zou je moeten doen om wél het goede genotype te krijgen?"]

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["Selectie"]
["\"Niets in de biologie heeft betekenis behalve in het licht van de evolutie\" (Theodosius Dobzhansky, 1973)"]
["Vul alleen gegevens in, in de groene velden. Wanneer je gegevens verandert in de andere cellen loop je kans dat de formules niet meer correct functioneren en daarmee de opdracht niet meer goed werkt."]
[]
["1. Een populatie is een groep dieren van dezelfde soort, die zich onderling (kunnen) voortplanten. Alle allelen die in een populatie voorkomen noem je de genenpoel. Streep door wat niet van toepassing is, en maak de zin af."]
[]
["Hoe meer allelen in de genenpoel, hoe",null,null,null,"kleiner / groter",null,"de variatie aan eigenschappen in de populatie, want"]
[]
["2. Kijk nog eens naar de zes ouderkippen (P) in je stamboek. Hoeveel verschillende allelen komen er voor in de genenpoel van deze (mini)populatie? Vul dat aantal in de tabel hieronder in. Tel ook het aantal allelen in de F1, de F2, de F3 en de F4."]
[null,null,null,"P","F1","F2","F3","F4"]
["aantal allelen in populatie"]
[]
["3. Waarschijnlijk ben je tijdens het fokken één of meerdere allelen kwijtgeraakt. Dat komt voor een deel doordat je ideale kip bepaalde eigenschappen juist niet moest hebben. Ben je ook allelen kwijtgeraakt van genen waar je eigenlijk helemaal niet op gelet hebt? (En zo ja, welke?)"]
[]
[]
["4. Streep door wat niet van toepassing is."]
[null,"Door fokken (kunstmatige selectie) neemt de variatie in de populatie ",null,null,null,null,null,null," toe  /  af."]
[null,"Door fokken (kunstmatige selectie) neemt de hoeveel homozygote genotypen",null,null,null,null,null,null,null,"  toe  /  af."]
[null,"In vergelijking met vroeger worden",null,null,null," meer  /  minder",null,"ouderdieren gebruikt voor de productie van een bepaalde hoeveelheid nakomelingen."]
[null,"Hoe minder ouderdieren je gebruikt, des te",null,null,null,null," meer  /  minder",null,"variatie je verliest."]
[null,"Hoe zeldzamer een bepaald allel, hoe",null,null,null,"sneller  /  minder snel",null,null,"dat allel uit de populatie verdwijnt."]
[]
["5: Hieronder staan alle kippen per generatie weergegeven die zijn gefokt onder jouw toezicht. Je hebt gezien dat je uiteindelijk na een aantal keer selecteren een bepaald genotype met bijbehorend fenotype kunt krijgen. Bekijk de allelen per generatie. Welke selectie stap (welke generatie) heeft het meest bijgedragen aan het eindresultaat? Schrijf je keuze (A / B / C / D of E) in het groene veld hieronder"]
[null,"A: In generatie P"]
[null,"B: In generatie F1"]
[null,"C: In generatie F2"]
[null,"D: In generatie F3"]
[null,"E: Dat kan ik niet zeggen op basis van hoe de gegevens hier zijn weergegeven"]
[]
[]
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["P",{"$f":"'1) Fok je ideale kip'!B50","v":"Bob"},null,{"$f":"'1) Fok je ideale kip'!D50","v":"Dirk"},null,{"$f":"'1) Fok je ideale kip'!F50","v":"Otto"},null,null,null,{"$f":"'1) Fok je ideale kip'!J50","v":"Anne"},null,{"$f":"'1) Fok je ideale kip'!L50","v":"Christina"},null,{"$f":"'1) Fok je ideale kip'!N50","v":"Puk"}]
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[]
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[]
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[]
["6: Door het percentage van ieder allel per generatie te berekenen kan je beter zien welke selectie stappen een grote invloed hadden op het eindproduct. Door deze gegevens in grafieken te zetten per gen krijg je helemaal een goed beeld wat er in iedere selectie stap is gebeurt. Bekijk de grafieken en kijk naar je antwoord op vraag 5. Klopte dit antwoord? Schrijf je antwoord helemaal onderaan de grafieken in het groene veld."]
[]
[]
[null,"Allel a / A"]
[null,"a",null,null,null,"A"]
[null,"generatie","totaal","procent",null,"generatie","totaal","procent"]
["P","P",{"$f":"SUM(AA33:AN33)","v":9},{"$f":"C77/0.12","v":75},null,"P",{"$f":"12-C77","v":3},{"$f":"G77/0.12","v":25}]
["F1000","F1",{"$f":"SUM(AA41:AN41)","v":0},{"$f":"C78/0.12","v":0},null,"F1",{"$f":"12-C78","v":12},{"$f":"G78/0.12","v":100}]
["F2000","F2",{"$f":"SUM(AA49:AN49)","v":0},0,null,"F2",{"$f":"12-C79","v":12},{"$f":"G79/0.12","v":100}]
["F3000","F3",{"$f":"SUM(AA57:AN57)","v":0},0,null,"F3",12,100]
["F4000","F4",{"$f":"SUM(AA65:AN65)","v":0},0,null,"F4",12,100]
[]
[]
[]
[]
[]
[]
[]
[]
[null,"Allel b / B","gen b / B"]
[null,"b","b",null,null,"B","B"]
[null,"generatie","totaal","procent",null,"generatie","totaal","procent"]
["P","P",{"$f":"SUM(AA34:AN34)","v":2},{"$f":"C93/0.12","v":16.666666666666668},null,"P",{"$f":"12-C93","v":10},{"$f":"G93/0.12","v":83.333333333333343}]
["F1000","F1",{"$f":"SUM(AA42:AN42)","v":0},{"$f":"C94/0.12","v":0},null,"F1",{"$f":"12-C94","v":12},{"$f":"G94/0.12","v":100}]
["F2000","F2",{"$f":"SUM(AA50:AN50)","v":0},0,null,"F2",{"$f":"12-C95","v":12},{"$f":"G95/0.12","v":100}]
["F3000","F3",{"$f":"SUM(AA58:AN58)","v":0},0,null,"F3",12,100]
["F4000","F4",{"$f":"SUM(AA66:AN66)","v":0},0,null,"F4",12,100]
[]
[]
[]
[]
[]
[]
[]
[]
[null,"Allel c / C","gen c / C"]
[null,"c","c",null,null,"C","C"]
[null,"generatie","totaal","procent",null,"generatie","totaal","procent"]
["P","P",{"$f":"SUM(AA35:AN35)","v":1},{"$f":"C109/0.12","v":8.3333333333333339},null,"P",{"$f":"12-C109","v":11},{"$f":"G109/0.12","v":91.666666666666671}]
["F1000","F1",{"$f":"SUM(AA43:AN43)","v":0},{"$f":"C110/0.12","v":0},null,"F1",{"$f":"12-C110","v":12},{"$f":"G110/0.12","v":100}]
["F2000","F2",{"$f":"SUM(AA51:AN51)","v":0},0,null,"F2",{"$f":"12-C111","v":12},{"$f":"G111/0.12","v":100}]
["F3000","F3",{"$f":"SUM(AA59:AN59)","v":0},0,null,"F3",12,100]
["F4000","F4",{"$f":"SUM(AA67:AN67)","v":0},0,null,"F4",12,100]
[]
[]
[]
[]
[]
[]
[]
[]
[null,"Allel d / D","gen d / D"]
[null,"d","d",null,null,"D","D"]
[null,"generatie","totaal","procent",null,"generatie","totaal","procent"]
["P","P",{"$f":"SUM(AA36:AN36)","v":9},{"$f":"C125/0.12","v":75},null,"P",{"$f":"12-C125","v":3},{"$f":"G125/0.12","v":25}]
["F1000","F1",{"$f":"SUM(AA44:AN44)","v":0},{"$f":"C126/0.12","v":0},null,"F1",{"$f":"12-C126","v":12},{"$f":"G126/0.12","v":100}]
["F2000","F2",{"$f":"SUM(AA52:AN52)","v":0},0,null,"F2",{"$f":"12-C127","v":12},{"$f":"G127/0.12","v":100}]
["F3000","F3",{"$f":"SUM(AA60:AN60)","v":0},0,null,"F3",12,100]
["F4000","F4",{"$f":"SUM(AA68:AN68)","v":0},0,null,"F4",12,100]
[]
[]
[]
[]
[]
[]
[]
[]
[null,"Allel e / E","gen e / E"]
[null,"e","e",null,null,"E","E"]
[null,"generatie","totaal","procent",null,"generatie","totaal","procent"]
["P","P",{"$f":"SUM(AA37:AN37)","v":7},{"$f":"C141/0.12","v":58.333333333333336},null,"P",{"$f":"12-C141","v":5},{"$f":"G141/0.12","v":41.666666666666671}]
["F1000","F1",{"$f":"SUM(AA45:AN45)","v":0},{"$f":"C142/0.12","v":0},null,"F1",{"$f":"12-C142","v":12},{"$f":"G142/0.12","v":100}]
["F2000","F2",{"$f":"SUM(AA53:AN53)","v":0},0,null,"F2",{"$f":"12-C143","v":12},{"$f":"G143/0.12","v":100}]
["F3000","F3",{"$f":"SUM(AA61:AN61)","v":0},0,null,"F3",12,100]
["F4000","F4",{"$f":"SUM(AA69:AN69)","v":0},0,null,"F4",12,100]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[null,"P"]
[null,"Dirk",null,"Puk"]
[null,"A","a","a","a"]
[null,"B","B","B","B",null,null,{"$f":"B202","v":"A"},null,{"$f":"C202","v":"a"}]
[null,"C","C","C","C",null,{"$f":"D202","v":"a"},null,null,{"$f":"IF(EXACT(UPPER(J203),J203),J203&G204,G204&J203)","v":"aa"}]
[null,"d","d","d","d"]
[null,"E","E","E","e",null,{"$f":"E202","v":"a"},{"$f":"IF(EXACT(UPPER(H203),H203),H203&G206,G206&H203)","v":"Aa"},null,{"$f":"IF(EXACT(UPPER(J203),J203),J203&G206,G206&J203)","v":"aa"}]

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["Natuurlijke selectie"]
["\"Niets in de biologie heeft betekenis behalve in het licht van de evolutie\" (Theodosius Dobzhansky, 1973)"]
["Vul alleen gegevens in, in de groene velden. Wanneer je gegevens verandert in de andere cellen loop je kans dat de formules niet meer correct functioneren en daarmee de opdracht niet meer goed werkt."]
[]
["Je hebt je ideale kip gefokt. Door te selecteren zijn er bepaalde genotypen met bijbehorende fenotypen ontstaan. In de natuur komen ook selectie stappen voor. Hieronder staan dezelfde grafieken als bij tabblad 2) Selectie, maar dan zit er tussen iedere meting 1000 generaties. In de volgende vragen ga je een scenario bedenken hoe jou kip door middel van natuurlijke selectie ook had kunnen ontstaan."]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
[]
["Allel A/a ","Dit gen codeert voor een groeihormoon, een eiwit dat onder andere zorgt voor de aanmaak van spierweefsel. De volgende allelen en fenotypen komen voor: "]
[null,"A: De kip heeft na 6 weken een lichaamsgewicht van 2,5 kg. Door de snelle groei heeft de kip gezondheidsproblemen en leeft niet langer dan een half jaar. "]
[null,"a: De kip bereikt na 14 weken een lichaamsgewicht van 2,5 kg. De kip kan onder normale omstandigheden vijf jaar of langer in leven blijven."]
[]
["Allel B/b","Dit gen codeert voor een hormoon dat invloed heeft op het sociaal gedrag van de kip. De volgende allelen en fenotypen komen voor: "]
[null,"B: De kip vertoont agressief gedrag (pikken) richting soortgenoten, met name wanneer de kippen dicht op elkaar zitten of wanneer de eigen kuikens worden aangevallen."]
[null,"b: De kip is relatief passief richting soortgenoten; de eigen kuikens worden nauwelijks beschermd tegen aanvallen van andere kippen."]
[]
["Allel C/c ","Dit gen codeert voor een enzym dat een rol speelt bij de pigmentatie (kleuring) van de veren. De volgende allelen en fenotypen komen voor:"]
[null,"C: De kip heeft egaal witte veren. "]
[null,"c: De kip heeft zwart-wit gespikkelde veren."]
[]
["Allel D/d ","Dit gen codeert voor een eiwit dat invloed heeft op de hoeveelheid energie die een kip investeert in voortplanting. De volgende allelen en fenotypen komen voor: "]
[null,"D: De kip investeert veel energie in voortplanting, en produceert gemiddeld 200 eieren per jaar. Hierdoor is de lichaamsgrootte van het dier beperkt."]
[null,"d: De kip investeert minder energie in voorplanting, en kan per jaar hooguit 50 eieren produceren. De eierproductie van het dier gaat niet ten koste van de lichaamsgroei."]
[]
["Alel E/e ","Dit gen codeert voor een enzym dat invloed heeft op de veergroei op de kop van de kip. De volgende allelen en fenotypen komen voor:"]
[null,"E: De kip heeft een groot aantal kuifveren op zijn kop. "]
[null,"e: De kip heeft geen kuifveren op zijn kop."]
[]
["1: jij hebt in tabblad 1 de ideale",null,null,null,{"$f":"'1) Fok je ideale kip'!B18","v":0},"geprobeerd te fokken. Je hebt geleerd dat een alles alleen behouden wordt in een populatie wanneer het allel geen nadeel oplevert."]
[null," Bekijk de verschillende allelen in de grafieken hierboven. Geef in de tabel hieronder aan welke genen zijn verdwenen."]
[]
[]
["2: Ging het verdwijnen geleidelijk of vrijwel ineens? Het plotseling verdwijnen van een allel heeft vaak een ramp als oorzaak. Geleidelijke verdwijning niet. Geef voor ieder allel aan of het verdwijnen plotseling ging of geleidelijk. Geef daarachter aan wat de reden hiervan geweest zou kunnen zijn, als je kijkt naar de functie van het betreffende gen."]
[]
[null,"allel","afgenomen / toegenomen / stabiel",null,null,"plotseling / geleidelijk",null,"functie",null,"mogelijke oorzaak"]
[null,"a",null,null,null,null,null,"groeihormoon"]
[null,"A"]
[null,"b",null,null,null,null,null,"gedrag"]
[null,"B"]
[null,"c",null,null,null,null,null,"veer kleur"]
[null,"C"]
[null,"d",null,null,null,null,null,"voortplanting"]
[null,"D"]
[null,"e",null,null,null,null,null,"kop veren"]
[null,"E"]
[]
["3: Je hebt geleerd dat een allel alleen meer voorkomt in een populatie wanneer het allel een reproductie voordeel geeft. Met andere woorden, door deze eigenschap komen er meer nakomelingen die ook weer nakomelingen krijgen ten opzichte van de dieren die niet dit allel hebben. Vaak is dit een geleidelijke toename van een allel, dus een verklaring voor de geleidelijke afname van het andere allel voor hetzelfde gen. Geef in de tabel hierboven aan voor ieder allel dat is toegenomen over de verschillende generaties wat het voordeel geweest zou kunnen zijn voor dit allel."]
[]
["4:  Sommige genen blijven stabiel binnen een populatie. Geef in bovenstaande tabel aan welke allelen stabiel zijn. Probeer hier een verklaring voor te geven."]
[]
[]
["5: Je hebt geleerd dat een definitie van een soort is (Nectar H6 \"Soorten en Populaties\" blz. 195, dat het geen vruchtbare nakomelingen kan krijgen wanneer het gekuist wordt met een ander soort. Bedenk een scenario voor de soortvorming van jou kip. Geef aan of het allopatrische soortvorming of sympatrische soortvorming betreft en of seksuele selectie een rol heeft gespeeld. Deze termen hebben we behandeld in paragraaf 7.2 \"ontstaan van nieuwe soorten\""]
[]
[]
[]
[]
["6: Gebruik de gegevens uit de tabel die je hebt gemaakt en het antwoord op vraag 4 om het evolutionaire verhaal te vertellen van jou kip. Maak er een vloeiend en boeiend verhaal van waarin je kort voor iedere periode / 1000 generaties beschrijft wat er gebeurt. Ga voor het gemak uit dat er tussen iedere 1000 generaties ongeveer evenveel jaren zitten. Generatie P leefde dus ongeveer 4000 jaar geleden als wilde kip.\nVerwerk de termen \"strugle voor life\" en \"survival of the fittest\" in je verhaal."]
[null,"omschrijving omgeving, dieren in P (genotype en fenotype) en manier van soortvorming"]
[]
[]
[]
[null,"omschrijving omgeving, dieren in F1000 (genotype en fenotype) en gebeurtenissen de afgelopen periode waardoor dit genotype is ontstaan"]
[]
[]
[]
[null,"omschrijving omgeving, dieren in F2000 (genotype en fenotype) en gebeurtenissen de afgelopen periode waardoor dit genotype is ontstaan"]
[]
[]
[]
[null,"omschrijving omgeving, dieren in F3000 (genotype en fenotype) en gebeurtenissen de afgelopen periode waardoor dit genotype is ontstaan"]
[]
[]
[]
[null,"omschrijving omgeving, dieren in F4000 (genotype en fenotype) en gebeurtenissen de afgelopen periode waardoor dit genotype is ontstaan"]
[]
[]
[]
["7. In de tabel hieronder worden kunstmatige selectie en natuurlijke selectie met elkaar vergeleken. verwijder in de groene velden wat niet van toepassing is."]
[]
[null,null,null,null,null,null,null,"natuurlijke selectie",null,null,null,"kunstmatige selectie"]
[null,"selectie door…",null,null,null,null,null,"de omgeving / de mens",null,null,null,"de omgeving / de mens"]
[null,"aantal eigenschappen waarop wordt geselecteerd ",null,null,null,null,null,"groot / klein",null,null,null,"groot / klein"]
[null,"gemiddeld aantal kinderen per ouder",null,null,null,null,null,"veel / weinig",null,null,null,"veel / weinig"]
[null,"verandering gaat...",null,null,null,null,null,"snel / langzaam",null,null,null,"snel / langzaam"]

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["Vorm en functie"]
["\"Niets in de biologie heeft betekenis behalve in het licht van de evolutie\" (Theodosius Dobzhansky, 1973)"]
["Vul alleen gegevens in, in de groene velden. Wanneer je gegevens verandert in de andere cellen loop je kans dat de formules niet meer correct functioneren en daarmee de opdracht niet meer goed werkt."]
[]
["Zoals je weet gaan vorm en functie samen binnen de biologie. In de simulatie van evolutie hebben we gekeken naar slechts 5 genen. In werkelijkheid zijn er natuurlijk veel meer genen betrokken. Neem als voorbeeld een giraf; wanneer een gen zorgt voor langere poten, zal een ander gen moeten zorgen voor een langere nek. Anders zal de giraf niet meer met zijn kop bij de grond kunnen om water te drinken. Een ander voorbeeld is de Tyrannosaurus Rex. Dit beest heeft een hele grote kop en loopt op twee poten. Daardoor heeft hij ook een grote staart nodig om in evenwicht te blijven. "]
[]
["1) Hieronder staat de tekst die je bij \"tabblad 3) Natuurlijke selectie\" hebt geschreven. (Deze tekst is hier niet meer aan te passen).  Lees de beschrijvingen die je hebt gemaakt hieronder en voeg een kleine omschrijving van het uiterlijk van de kip weer onder iedere generatie. Probeer de omschrijving te koppelen aan de allelen die je eerder hebt omschreven. "]
[null,{"$f":"'3) Natuurlijke selectie'!B70","v":"omschrijving omgeving, dieren in P (genotype en fenotype) en manier van soortvorming"}]
[]
[]
[]
[]
[]
[]
[null,{"$f":"'3) Natuurlijke selectie'!B74","v":"omschrijving omgeving, dieren in F1000 (genotype en fenotype) en gebeurtenissen de afgelopen periode waardoor dit genotype is ontstaan"}]
[]
[]
[]
[]
[]
[]
[null,{"$f":"'3) Natuurlijke selectie'!B78","v":"omschrijving omgeving, dieren in F2000 (genotype en fenotype) en gebeurtenissen de afgelopen periode waardoor dit genotype is ontstaan"}]
[]
[]
[]
[]
[]
[]
[null,{"$f":"'3) Natuurlijke selectie'!B82","v":"omschrijving omgeving, dieren in F3000 (genotype en fenotype) en gebeurtenissen de afgelopen periode waardoor dit genotype is ontstaan"}]
[]
[]
[]
[]
[]
[]
[null,{"$f":"'3) Natuurlijke selectie'!B86","v":"omschrijving omgeving, dieren in F4000 (genotype en fenotype) en gebeurtenissen de afgelopen periode waardoor dit genotype is ontstaan"}]
[]
[]
[]
[]
[]
[]
[]
["2: Sommige aanpassingen die in het verleden zijn ontstaan, zijn in de \"nieuwe versie\" niet meer nodig. Toch verdwijnt zo'n aanpassing niet, omdat het ook geen nadeel is om deze aanpassing te hebben. Een voorbeeld is het staartbeen bij de mens. Wij hebben geen staart meer, maar hebben toch nog de botjes ervoor in ons genetisch bouwplan en dus in ons lichaam zitten. Dit soort \"organen\" die we nog wel hebben maar niet meer nodig hebben heten rudimentaire organen. (zie ook paragraf 7.3 \"ontstaan van nieuwe soorten\") Geef 2 voorbeelden van rudimentaire organen in de F4000. Geef ook aan waar ze vroeger voor dienden en in welke F ze nog wel een functie hadden."]
[]
[]
[]
[]
["3: Teken voor iedere generatie uit de opdracht hoe de kip er ongeveer uitgezien zou hebben. Scan de tekening of maak een foto en plak die hieronder in dit bestand. Geef met pijltjes en woorden de belangrijkste uiterlijke kenmerken aan. Doe dit ook met de organen die later rudimentair zijn geworden."]

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["Nieuwe variatie"]
["\"Niets in de biologie heeft betekenis behalve in het licht van de evolutie\" (Theodosius Dobzhansky, 1973)"]
["Vul alleen gegevens in, in de groene velden. Wanneer je gegevens verandert in de andere cellen loop je kans dat de formules niet meer correct functioneren en daarmee de opdracht niet meer goed werkt."]
[]
["In tabblad 5) \"nieuwe variatie\" heb je gezien dat er door een mutatie, insertie of deletie een nieuwe variant van een gen (een allel) kan ontstaan. Ook heb je gezien dat mutaties maar weinig voorkomen. \nHoe kan zo'n nieuw allel zich dan toch verspreiden in een populatie? Daar gaan we hier naar kijken. "]
[]
["1: Door fokken (kunstmatige selectie) neemt de hoeveelheid variatie in de populatie af. Ook in de natuur verdwijnen voortdurend allelen. Namelijk door natuurlijke selectie. Allelen die coderen voor eigenschappen die ongunstig zijn voor de soort worden door natuurlijke selectie weggefilterd. Hieronder zie je de allelen van de \"oerkip\". Bekijk het genotype en vergelijk dit met het genotype van jouw kippen uit de F4 van tabblad \"1) Fok je ideale kip\". Is het nog mogelijk om deze oerkip terug te fokken? Leg uit welke allelen er zijn verdwenen."]
[]
[]
[null,"Oerkip"]
["gen1","a","a"]
["gen2","B","B"]
["gen3","C","C"]
["gen4","d","d"]
["gen5","e","e"]
[]
["2a: Wanneer allelen verdwenen zijn in een populatie wil dat niet zeggen dat niet altijd zeggen dat deze allelen definitief verdwenen zijn uit die populatie. Door migratie van ellelen van de ene naar de andere populatie kan de allel frequentie veranderen. Er zijn verschillende voorplantings scenario's denkbaar die allemaal een effect op de genetische diversiteit van een populatie hebben. Vul onderstaande tabel in."]
[null,null,null,null,null,null,"andere soort",null,null,null,null,null,null,null,"zelfde soort"]
[null,null,null,null,null,null,null,null,null,null,null,null,null,null,"ander ras",null,null,null,"zelfde ras"]
[null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,"ander populatie",null,"zelfde populatie"]
[null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,null,"ander gezin","zelfde gezin"]
[]
[null,"F1 levend (Ja / Nee)"]
[null,"F1 gezond (Ja / Nee)"]
[null,"genetische diversiteit populatie (neem toe / neemt af)"]
[]
["2b: Kijk goed naar de opbouw van de tabel. Bekijk het genotype van de oerkip en van jouw populatie kippen. Teken een cladogram waarin je de oerkip, vleeskip, legkip en dierkip weergeeft. Maak een foto van je tekening en plak hem hieroner in het groene veld."]
[]
[]
["2c: Leg uit dat je met behulp van DNA analyse een cladogram nog nauwkeurig kan samenstellen."]
[]
[]
["3: Wanneer er alleen in de loop der tijd door selectie alleen maar allelen verloren gaan, zou al het leven op een gegeven moment uitsterven. Probeer deze stelling uit te leggen in het perspectief van evolutie."]
[]
[]
["4: Gelukkig kan er ook nieuwe variatie in DNA ontstaan. Twee processen die zorgen voor nieuwe variatie zijn mutatie en recombinatie"]
[]
[]
["Mutatie ","Mutatie is het optreden van veranderingen in de basenvolgorde van het DNA. Mutaties ontstaan spontaan, of worden veroorzaakt door straling of bepaalde (mutagene) stoffen. De meeste mutaties hebben óf geen enkel effect, óf zorgen ervoor dat een gen niet meer goed functioneert."]
["Re- combinatie ","De meeste dieren zijn diploïd: in elke cel komen van ieder gen of chromosoom twee exemplaren voor (één van vader en één van moeder). Bij geslachtelijke voortplanting ontstaan steeds nieuwe combinaties van allelen. Dat noem je recombinatie. Veel eigenschappen worden door meer dan één enkel gen bepaald. Hierdoor kunnen door recombinatie ook nieuwe eigenschappen ontstaan. "]
[]
["Maak het onderstaande schema compleet. Noteer de begrippen ‘mutatie’ en ‘recombinatie’ in het juiste vak. Geef bij alle pijlen met een plus (+) of een min (-) aan of er sprake is van een positief of een negatief verband."]
[]
[null,null,null,null,"\"+/-\"",null,"aantal allelen in de genenpoel"]
[]
[null,null,null,null,null,null,null,null,"\"+/-\""]
[]
[null,null,null,null,"\"+/-\"",null,"variatie in genotypen in de populatie"]
[]
[null,null,null,null,null,null,null,null,"\"+/-\""]
[]
[null,null,null,null,null,null,"variatie in fenotypen in de populatie"]
[]
[]
["5. Leg uit dat fokken niet alleen een vorm van selectie, maar ook een vorm van recombinatie is. "]
[]
[]
["6: Leg uit wat er gebeurt op aminozuur niveau bij een mutatie, deletie en insertie. Gebruik in je antwoorden de woorden DNA triplet en codon."]
[null,"mutatie (substitutie):"]
[null,"deletie:"]
[null,"insertie:"]
[]
["7: Leg uit hoe een verandering op aminozuurniveau een verandering in eiwit kan geven"]
[]
[]
["8: Leg uit hoe een verandering in eiwit een verandering in fenotype kan geven"]
[]
[]
["9: Een mutatie, deletie of insertie kunnen invloed hebben op de vouwing van een eiwit in een organisme. Deze foutjes ontstaan vaak min of meer willekeurig in het genoom. Toch zijn er genen waar geen enkele mutatie in voorkomt. Leg uit hoe dit komt?"]
[]
[]
["10: In andere genen komen relatief veel variaties voor. Hieronder zie je een mutatie simulatie voor het gen a van de oerkip in een populatie van 1000 kippen in de F1. Pas het getal achter het veld \"mutatie frequentie\" aan en kijk wat er gebeurt. Leg uit dat met een lage mutatie frequentie, zonder recombinatie de kans op het genotype AA uiterst klein is."]
[]
[]
["mutatie frequentie",null,0.01]
["gevonden mutaties F1:",null,{"$f":"E84+(F84*2)","v":13}]
[]
[null,"gameten",null,null,null,null,"gameten"]
[null,"a","a",null,null,null,"a","a"]
["mutatie","A",null,null,null,"mutatie","A"]
["genotype ",{"$f":"B79&C79","v":"aa"},null,null,null,"genotype ",{"$f":"G79&H79","v":"aa"}]
[]
[null,"totale populatie",null,"aa","Aa","AA"]
["aantal:",{"$f":"SUM(D84:F84)","v":1000},null,{"$f":"SUM(Q91:Q1090)","v":987},{"$f":"SUM(R91:R1090)","v":13},0]
["aantal %",{"$f":"B84/10","v":100},null,{"$f":"D84/10","v":98.7},1.3,0]
[]
[]
[]
[null,"P (moeder)",null,null,null,null,"P (vader)",null,null,null,null,"F1 (kind)"]
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["Hardy-Weinberg-evenwicht"]
["\"Niets in de biologie heeft betekenis behalve in het licht van de evolutie\" (Theodosius Dobzhansky, 1973)"]
["vul alleen gegevens in, in de groene velden. Wanneer je gegevens verandert in de andere cellen loop je kans dat de formules niet meer correct functioneren en daarmee de opdracht niet meer goed werkt."]
[]
["Er zijn mensen die stellen dat evolutie niet bestaat. Om geen evolutie plaats te laten vinden, zouden er specifieke omstandigheden moeten plaatsvinden, waardoor er geen verandering in de genenpool plaatsvindt.\nMet andere woorden; de mate waarin ieder allel aanwezig is in een populatie mag niet veranderen. Wanneer dit het geval is, spreekt met van het Hardy-Weinberg evenwicht. Dit evenwicht is te berekenen met een formule waarbij je de allelfrequentie kunt invullen.\nEr zijn twee formules die elkaars afgeleide zijn: p² + 2pq + q² = 1 en p+q = 1.\nHierbij is p het dominante allel en q het recessieve allel. \nHieronder staat een stappenplan dat je helpt te leren rekenen met allelen in een populatie.\nWe gaan nu kijken of de allelfrequentie is veranderd voor jouw populatie kippen."]
[]
["1: Een paleontoloog heeft in een gebied zes fossielen gevonden van kuikens in een nest. De kuikens lijken zo'n 4 weken oud te zijn. Opvallend is dat er 1 exemplaar tussen zit, die behoorlijk veel groter is dan de rest. Op welke twee manieren kan de leeftijd bepaald worden van deze fossielen?"]
["manier 1:"]
["manier 2:"]
[]
["2: De fossielen blijken zo'n 4000 jaar oud te zijn. Uit andere opgravingen blijkt dat de totale populatie ongeveer uit 1200 kippen heeft bestaan. Op basis van deze 2 bevindingen stellen de onderzoekers dat waarschijnlijk 1/6 deel van de populatie kippen al na  6 weken na de geboorte volgroeid was in tegenstelling tot de andere kippen. De onderzoekers denken dat dit met Gen A/a te maken heeft: Dit gen codeert voor een groeihormoon, een eiwit dat onder andere zorgt voor de aanmaak van spierweefsel. Kippen met het dominante allel A zijn na 6 weken al volgroeid.\na) \tBereken de allelfrequenties voor A en a \nb) Bbereken alle genotypefrequenties."]
[]
["Bij monogene overerving wordt er gekeken naar 1 gen met 2 allelen. Hieronder staat een uitgewerkt voorbeeld voor monogene overerving. Links staat het stappenplan, rechts staat een uitgewerkt voorbeeld met behulp van dit stappenplan."]
[]
["Stappen plan:"]
["stap 1: Lees , onderstreep en schrijf op wat je weet",null,null,null,null,null,null,null,null,null,null,null,null,null,"P","q"]
["Lees de vraag nauwkeurig en onderstreep belangrijke informatie.\n",null,null,null,null,null,null,null,null,"P² + 2Pq + q² = 1\n",null,null,null,"P","P²","pq"]
["Schrijf de Hardy-Weinberg formules op en maak de bijbehorende tabel. Dit je het gereedschap!",null,null,null,null,null,null,null,null,"P + q = 1",null,null,null,"q","pq","q²"]
["P is het dominante gen, q recessief"]
[]
["stap 2: Bereken de homozygoot recessieve genotypefrequentie (q²)\n",null,null,null,null,null,null,null,null,"200 = AA / Aa",null,null,null,null,"P","q"]
["Start altijd met het berekenen van q². ",null,null,null,null,null,null,null,null,"aa = 1200-200",null,null,null,"P"]
["Deze groep bestaat uit homozygoot recessief en dus hoort er maar 1 fenotype bij dit genotype.",null,null,null,null,null,null,null,null,"aa =1000",null,null,null,"q",null,0.83]
["Schrijf q² altijd als een fractie van 1",null,null,null,null,null,null,null,null,"aa = qq"]
["Zet het getal in de tabel",null,null,null,null,null,null,null,null,"1000 / 12.00 ≈  83%"]
[]
["stap 3: Bereken de recessieve allel frequentie (q)",null,null,null,null,null,null,null,null,"q² = 0.83",null,null,null,null,"P",0.91]
["Door de wortel van q² te nemen krijg je q",null,null,null,null,null,null,null,null,"q = √0.88",null,null,null,"P"]
[null,null,null,null,null,null,null,null,null,"q ≈ 0.91",null,null,null,0.91,null,{"$f":"P23","v":0.83}]
[]
["stap 4: Bereken de allelfrequentie van het dominante allel (P)",null,null,null,null,null,null,null,null,"P + q = 1",null,null,null,null,0.09,{"$f":"P27","v":0.91}]
["Met behulp van de formule P + q = 1 kun je nu P berekenen.",null,null,null,null,null,null,null,null,"P + 0.91 = 1",null,null,null,0.09]
["Nu zijn alle allel frequenties bekend. ",null,null,null,null,null,null,null,null,"P = 0.09",null,null,null,{"$f":"N29","v":0.91},null,{"$f":"P23","v":0.83}]
[]
["stap 5: Bereken de overige genotypefrequenties",null,null,null,null,null,null,null,null,"P² = 0.09 x 0.09",null,null,null,null,{"$f":"O31","v":0.09},{"$f":"P31","v":0.91}]
["Nu P en q zijn uitgerekend is het mogelijk om de genotypefrequenties te berekenen.",null,null,null,null,null,null,null,null,"P² ≈ 0.008",null,null,null,{"$f":"N32","v":0.09},8.0000000000000002E-3,0.08]
[null,null,null,null,null,null,null,null,null,"Pq = 0.91 x 0.09",null,null,null,{"$f":"N33","v":0.91},0.08,{"$f":"P33","v":0.83}]
[null,null,null,null,null,null,null,null,null,"Pq ≈ 0.08"]
[]
["stap 6: Geef antwoord op de vraag"]
["Kijk goed wat er precies van je wordt gevraagd en let op dat je de juiste allel- en gennamen gebruikt."]
[]
["2a:"]
["2b:"]
[]
["3a: Vaak gaan allelen na een lange periode naar fixatie. Voor het gemak gaan we er nu even vanuit dat dit niet het geval was. \nWanneer er geen evolutie zou hebben plaatsgevonden de afgelopen 4000 jaar, zou dus de allelfrequentie en genotypefrequentie onveranderd moeten zijn. Om dit te controleren, kijken de onderzoekers naar de huidige populatie kippen in het gebied. Er blijken nu 3495 kippen in het gebied te lopen, waarvan 3026 kippen pas na 14 weken volgroeid waren. Bereken de allelfrequentie en de genotypefrequentie van de huidige populatie kippen. Gebruik hiervoor de zes stappen, zoals je hierboven hebt geleerd. Vul onderstaande tabel in."]
[]
[null,null,"P","q"]
[null,"P","P²","pq"]
[null,"q","pq","q²"]
[]
["3b: allel frequentie:"]
["3c: genotypefrequentie:"]
[]
["4: Vergelijk de allelfrequentie tussen de kippenpopulatie 4000 jaar geleden en de huidige kippen populatie. Heeft er evolutie plaatsgevonden? Leg je antwoord uit. "]
[]
[]
["5: Wanneer er voor dit specifieke gen geen verandering in allel- en genotypefrequentie had plaatsgevonden, is dit dan sluitend bewijs dat er geen evolutie heeft plaatsgevonden?"]

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["Fok evolutie"]
["\"Niets in de biologie heeft betekenis behalve in het licht van de evolutie\" (Theodosius Dobzhansky, 1973)"]
["vul alleen gegevens in, in de groene velden. Wanneer je gegevens verandert in de andere cellen loop je kans dat de formules niet meer correct functioneren en daarmee de opdracht niet meer goed werkt."]
[]
["Je hebt nu ook berekend dat er een verschuiving in de allelfrequentie van de populatie zit. De vraag is nu waardoor er geen Hardy-Weinberg evenwicht is en waardoor er dus evolutie heeft plaatsgevonden. \nEr zijn in totaal 5 voorwaarden waaraan een populatie moet voldoen om in een Hardy-Weinberg evenwicht te verkeren. Wanneer één of meerdere van deze voorwaarden niet kloppen, zal er een verschuiving in de allelfrequentie plaatsvinden en zal er dus evolutie plaatsvinden. Deze voorwaarden zijn:\n1) Er mogen geen nieuwe mutaties ontstaan\n2) De partner keuze is willekeurig\n3) Er vindt geen selectie plaats\n4) Er is geen geneflow\n5) Er is sprake van een extreem grote populatie\nHieronder gaan we deze voorwaarden een voor een onder de loep nemen."]
[]
["1: Er mogen geen nieuwe mutaties bij komen. Wanneer er nieuwe mutaties in een populatie ontstaan heeft dit effect op de genenpool. Hierdoor kan er bijvoorbeeld een heterozygoot genotype ontstaan. \nBekijk de genenpool van de F1. Bij welke allelen is er (waarschijnlijk) sprake geweest van een mutatie? Leg je antwoord uit."]
[]
[]
["2: Er vindt willekeurig paring plaats. Wanneer individuen een voorkeur hebben voor voortplanten, ontstaat er een shift in de genenpool. \nBekijk nog een keer de verandering van allelen over de verschillende generaties in tabblad 2. Heeft er willekeurige paring plaatsgevonden?\n"]
[]
[]
["3: Er vindt geen natuurlijke selectie plaats. Wanneer dieren met een bepaald genotype (dus fenotype) korter leven of voor meer nakomelingen zorgen, heeft dit een effect op de genpool.\nBekijk nog eens het verhaal dat je hebt geschreven. Heeft er ergens in het evolutie scenario dat je hebt bedacht een reproductievoordeel of -nadeel plaatsgevonden?\n"]
[]
[]
["4: Er is geen geneflow. Met geneflow wordt bedoeld dat er een allel in of uit de populatie gaat, doordat er individuen van de ene naar de andere populatie gaan. Is er sprake geweest van geneflow bij de totstandkoming van jouw kip? Leg je antwoord uit.\n"]
[]
[]
["5: Er is sprake van een extreem grote populatie. Bij een kleine populatie vindt genetic drift plaats.\nEr zijn verschillende omstandigheden die tot genetic drif kunnen leiden die een groot effect kunnen hebben op de populatie. Twee voorbeelden hiervan zijn het  “bottleneck effect” en het “founder effect”.\n5a: Het bottleneck effect: Een  plotselinge verandering in de omgeving kan een grote invloed hebben op de grootte van de populatie. Bijvoorbeeld door een ramp kan de variatie in een genenpool ineens teruglopen, waardoor bijvoorbeeld bepaalde allelen verdwijnen.\nIs er van een botleneck effect sprake geweest bij de totstandkoming van jouw kip? Leg je antwoord uit."]
[]
[]
["5b: Het founder effect: Wanneer een aantal indiviuen afgezonderd raken van de grotere populatie en zich daarna voortplanten, heet dit het “founder effect”. Een relatief zeldzame mutatie kan na verloop van tijd dan aanwezig zijn in een groot deel van de populatie. \nIs er van een founder effect sprake geweest bij de totstandkoming van jouw kip? Leg je antwoord uit. \n"]
[]
[]
["5c: Hieronder zie je een simulatie model voor genetic drift. De partnerkeuze in generatie P kan een groot effect hebben op de verspreiding van een allel binnen de populatie na bijvoorbeeld 100 generaties.\nVerander het toeval getal en bekijk wat er met de allelfrequentie en genotypes gebeurt. Klik een aantal keer op de knop \"verander toeval getal\". Leg in eigen woorden uit wat er gebeurt en waarom."]
[]
[]
[null,null,null,"A","a"]
[null,null,null,0.09,0.91,"volgende generatie"]
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[null,"a",0.91,{"$f":"C31*D29","v":8.1900000000000001E-2},{"$f":"C31*E29","v":0.82810000000000006},{"$f":"SUM(D31:E31)","v":0.91}]
[null,"volgende generatie",null,{"$f":"SUM(D30:D31)","v":0.09},{"$f":"SUM(E30:E31)","v":0.91}]
[]
[]
[]
[]
[]
[]
[null,null,{"$f":"RAND()","v":0.41323802088637063}]
[]
[]
[]
[null,null,{"$f":"RAND()","v":0.79834390028442648}]
[]
[]
[]
[]
[null,null,{"$f":"RAND()","v":0.32665944169885053}]
[]
[]
["toeval getal",null,{"$f":"RAND()","v":0.22415301044090907}]
["standaard afwijking",null,5.0000000000000001E-3]
[]
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